A Bryological Online Journal
Frahmia is a "paperless journal" which is published in the internet. It was founded with the intention and attitude that always drove the late Jan-Peter Frahm to provide the means for bryologists to publish the results. Insofar it might be understood as the successor of the "Archive of Bryology" that was created by the late Jan-Peter Frahm in 1994.

It is likewise an open access journal distributed under the terms of the Creative Commons Attribution License 3.0 (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.

Issued Frahmia Volumes
  • Volume 1 (June 28, 2014)
    Schwarz, U. (2014). Personal Memories to Jan-Peter Frahm (1945 – 2014). Frahmia 1:1-3.
    Abstract: The author summarizes his very own memories to Jan-Peter Frahm reaching from the late 1980s until today. It also provides an insight why Frahmia, as a new bryological online journal was created..
  • Volume 2 (June 28, 2014)
    Schwarz, U. (2014). Contribution to the Bryophyte Flora of Shinagawa, Tokyo, Japan (日本関東地方東京都区部品川区のコケ). Frahmia 2:1-29.
    Abstract: The bryophyte flora of Shinagawa, Tokyo, Japan was investigated in 2008 and 2009. A detailed list consisting 62 genera with 94 species is given. Bruchia flexuosa is reported for the second time in Japan.
  • Volume 3 (July 19, 2014)
    Schwarz, U. (2014). Fissidens brevinervis Broth. – New to the Indian Moos Flora. Frahmia 3:1-6.
    Abstract: Fissidens brevinervis Broth. was found in Bangalore, Karnataka, India in 2013. It’s the first record for India and extends the distribution range of this species that was only known so far from China and Indonesia. A brief summary of the current status of the bryophyte flora of Bangalore and the Fissidens species of India are provided.
  • Volume 4 (July 25, 2014)
    Bansal, P.; Nath, V. (2014). Genus Bryum Hedw. in Peninsular India. Frahmia 4:1-11.
    Abstract: The present study is aimed mainly to document the diversity of genus Bryum Hedw. in Peninsular India. During the revision of the genus 26 taxa are distributed in four states (Karnataka, Kerala, Maharashtra, Tamil Nadu) and one union territory (Goa) of Western Ghats, whereas 8 species reported from 3 states (Andhra Pradesh, parts of Odisha and Tamil Nadu) of Eastern Ghats. Bryum tuberosum Mohamed & Damanhuri is reported for the first time from Shimoga district of Karnataka and B. wightii Mitt. from Munnar (Idukki district) of Kerala.
  • Volume 5 (August 03, 2014)
    Rajesh, K.P.; Manju, C.N. (2014). Bryophyte Diversity of the lowlands and midlands of Kozhikode District, Kerala, India. Frahmia 5:1-10.
    Abstract: This article presents the diversity of Bryophytes of the lowlands and midlands of the Calicut (Kozhikode) district of Kerala in the Peninsular India. It provides a list of 57 species, mainly from the thickly populated areas.
  • Volume 6 (August 11, 2014)
    Mufeed, B.; Deepa, K.M.; Haripriya, P.; Manju, C.N. (2014). Moss (Bryophyta) diversity of Thusharagiri Hills, Kerala, India. Frahmia 6:1-9.
    Abstract: This article presents the Moss (Bryophyta) diversity of Thusharagiri hills in the Western Ghats of Kozhikode district, Kerala, India. A total of 32 species were recorded from a small evergreen patch.
  • Volume 7 (September 9, 2014)
    Alam, A. (2014). Morphotaxonomy of Fossombronia Raddi (Metzgeriales) from Nilgiri hills (India). Frahmia 7:1-12.
    Abstract: The present study is intended primarily to report the diversity of genus Fossombronia Raddi in Nilgiri hills (India). Only 4 species, Fossombronia cristula Austin, F. himalayensis Kashyap, F. pusilla (L.) Dumort., and F. wondraczekii (Corda) Dumort. ex Lindb. have been collected during the recent investigation. Earlier describe species, F. foreaui Udar et Srivastava, from these hills was not found in the few recent collections hence its details is not included in this study.
  • Volume 8 (September 10, 2014)
    Rawat, K.K.; Verma, P.V. (2014). On a collection of liverworts from Tawang, Arunachal Pradesh, India with Frullania rhystocolea Herzog new to India. Frahmia 8:1-7.
    Abstract: During a small trip in 2002, a total of 24 species of liverworts, both terrestrial and epiphytic, were collected from Tawang and are being described here including Frullania rhystocolea Herzog as new to India and Blasia pusilla L. as new to Arunachal Pradesh, making the first ever recorded account of liverworts of Tawang, Arunachal Pradesh.
  • Volume 9 (October 8, 2014)
    Alam, A. (2014). Professor S. C. Srivastava- A Living Legend of Indian Bryology. Frahmia 9:1-10.
    Abstract: This article is an effort to admire the attainments of Professor Suresh Chandra Srivastava in the field of Indian Bryology.
  • Volume 10 (November 23, 2014)
    Schwarz, U. (2014). Bruchiaceae collected in Karnataka, India, with a synopsis of the family in India. Frahmia 10:1-9.
    Abstract: During the exploration of the bryophyte flora of Karnataka, India, 2 species belonging to the family of Bruchiaceae, Trematodon longicollis and Trematodon schmidii, were found. Both species are illustrated. An overview about all the 10 species of Bruchiaceae in India, inclusive a key, short description and line drawings are provided
  • Volume 11 (May 6, 2015)
    Deepa, K.M.; Manju, C.N.; Nair, Maya C.; Rajesh, K.P. (2015). Exormotheca ceylonensis Meijer (Marchantiophyta) - an endangered liverwort of India, now recorded from a highly threatened habitat of Kerala. Frahmia 11:1-4.
    Abstract: Exormotheca ceylonensis, a critically endangered species is reported from Kerala and is described with Photograps.
  • Volume 12 (April 22, 2016)
    Silva, J.B.; Pôrto, K.C. (2016). Can we use the acrocarpous moss gametophyte length to assess microclimatic conditions in harsh environmental?. Frahmia 12:1-15.
    Abstract: Bryophytes are small non-vascular plants which are very sensitive to narrow variations in environmental conditions and can be considered to be useful bioindicators of local environmental conditions. Althogh their growth is encouraged by constant wetness, stem growth is a vigorous measure which can work as a good proxy for local microclimatic conditions. This is important when faced with the predicted climate change in the Caatinga. Our aim is to validate a trait which is easy to measure and which can act as a local condition proxy in monitoring changes in harsh environments. Based on the Generalized Linear Model (GLM), gametophyte length vs. soil depth, and descriptive statistics, we evaluated the vigor of length measurements as a proxy for local conditions in harsh environments. Soil depth varied as expected for both shallow and deep soil islands. Gametophyte’s length is a trait with high variability in the moss under study and such variation is not correlated with soil depth. The distribution of length values was more aggregated in Campylopus pilifer, with stem length being higher than that seen in the literature for most of the selected gametophytes. We found that length is a poor predictor trait of local conditions in harsh environments. However, our results suggest that soil island can act as proof of harsh environments given that C. pilifer, a pioneer moss, possessed stems which were higher than expected
  • Volume 13 (August 28, 2016)
    Schwarz, U. (2016). Physcomitrium schummii sp. nov. from Karnataka, India with a synopsis of the Funariaceae in India. Frahmia 13:1-19.
    Abstract: During the exploration of the bryoflora of Karnataka, India one new species of Physcomitrium was discovered and described as new species. Furthermore a synopsis of the family in India, represented by 4 genera, including 13 species of Entosthodon, 7 species of Funaria, 1 species of Loiseaubryum and 12 species of Physcomitrium is given. Funaria excurrentinervis, F. sinuatolimbata, F. subimmarginata, and F. pulchra are transferred to the genus Entosthodon.
  • Volume 14 (January 5, 2017)
    Rawat, K.K; Sahu, V.; Singh, C.P.; Verma, P.K. (2017). Additions to the Bryophyte flora of Tawang, Arunachal Pradesh, India. Frahmia 14:1-17.
    Abstract: A total of 30 taxa of bryophytes are reported for the first time from Tawang district of Arunachal Pradesh, India, including 10 taxa as new to Arunachal Pradesh.
  • Volume 15 (November 5, 2017)
    Homm, Th. (2017). Leaf-born gemmae in Syntrichia virescens (De Not.) Ochyra - a neglected feature in bryological literature. Frahmia 15:1-4.
    Abstract: Leaf-born gemmae have been detected in specimens of Syntrichia virescens from Northwest Germany. This feature is not mentioned or even denied to exist in most of the recent European bryological literature. This paper provides a short description and illustrations of the gemmae.
  • Volume 16 (October 20, 2019)
    Schwarz U.; Schumm F. (2019). Contribution to the bryophyte flora of Madeira Island. Frahmia 16:1-23.
    Abstract: During field trips in Madeira 2018, 173 bryophyte taxa were collected. The list of the taxa and the location of collection is given. Platyhypnidium torrenticola, so far only know from the Canary Islands, was also found. An illustration and comments on this peculiar plant are provided.
  • Volume 17 (August 30, 2020)
    Gradstein, S.R. (2020). Checklist of the Liverworts and Hornworts of Ecuador. Frahmia 17:1-40.
    Abstract: 706 species of liverworts and 16 of hornworts are reported from Ecuador; a further twenty species are listed as doubtful. Endemism in the country is about 6%. For each accepted species, distribution within the country (by province) and elevational range is given. The disproportionate numbers of provincial records indicate that the exploration of the country is still incomplete, with exception perhaps of the Galápagos Islands.
  • Volume 18 (November 4, 2020)
    Schwarz, U. (2020). Pseudotaxiphyllum annii sp. nov., a new species of Plagiotheciaceae from Philippines. Frahmia 18:1-8.
    Abstract: During the exploration of the bryoflora of the Philippines one new, propagule bearing species of Pseudotaxiphyllum was discovered. The taxon is described and illustrated as P. annii. The relation to other propagule bearing taxa of the P. pohliaecarpum – complex is discussed. The critical review of recent works on Psuedotaxiphyllum, together with the current study led to a species concept based on propagule morphology, recognizing 6 species in East and Southeast Asia.
  • Volume 19 (December 2, 2020)
    Winter, G.; Schäfer-Verwimp, A. (2020). Re-evaluation of the taxonomic status of Frullania caulisequa and Frullania obcordata (Frullaniaceae, Marchantiophyta). Frahmia 19:1-21.
    Abstract: Jungermannia obcordata Lehm. & Lindenb. was published 1834 by Lehmann and transferred to Frullania in Synopsis Hepaticarum 1845. Yuzawa (1988) transferred 4 species as synonyms sub Frullania caulisequa (Nees) Mont., among them Frullania obcordata (Lehm. & Lindenb.) Lehm. & Lindenb. probably without having seen the type of Frullania caulisequa. The examinations of the types of Frullania obcordata (hb GLM) and Frullania caulisequa (hb STR) clearly proved both to be different. All specimens named since 1988 probably belong to Frullania obcordata, not to Frullania caulisequa and have to be re-checked. New synonyms of both species are provided and all synonyms known to us are listed. The lobuli of Frullania caulisequa indicate that it is close eto Frullania curvilobula Schäf.-Verw., D.F.Peralta & S.M.Siqueira. For F. caulisequa and allied species (“Diastaloba IV”) the new subgenus Caulisequa is proposed.
  • Volume 20 (December 20, 2020)
    Schäfer-Verwimp, A.; Winter, G. (2020). Frullania dasyueshanensis, a new species with terete perianth from Taiwan (Marchantiophyta: Frullaniaceae). Frahmia 20:1-14.
    Abstract: Frullania dasyueshanensis sp. nov. is described from Dasyueshan National Forest Recreation area and other localities in Taiwan. The new species was collected in mountain rainforests between 2230 and 2650 meter. It differs from all known Frullania species in Taiwan by its clavate-cylindrical terete perianth and can be clearly distinguished from the similar Frullania bhutanensis S.Hatt. in its overall smaller size, the large size of the lobules compared to the lobes and and the rounded, not appendiculate dorsal base of leaf lobe; Frullania sphaerantha S.Hatt. from India is best distinguished from the new species by smaller lobules compared with the lobes and the widely pyriform perianth with short beak. Frullania sinosphaerantha S.Hatt. et P.J.Lin is considered to be synonymous with Frullania bhutanensis. A key for Asian Frullania species with terete perianths is provided.
  • Volume 21 (January 26, 2021)
    Schwarz, U. (2021). On a very small collection of bryophytes from North Korea. Frahmia 21:1-4.
    Abstract: During a sightseeing trip to North Korea the author was able to collect a few bryophyte samples. Altogether 14 mosses and 1 liverwort was discovered at 2 collection sites.
  • Volume 22 (March 15, 2021)
    Frahm, J.-P.; Pócs, T. (ed.) (2021). East African Bryophytes XXXII. Campylopoideae Records. Frahmia 22:1-5.
    Abstract: More then 30 years ago, J.-P. Frahm prepared a manuscript about African Campylopoideae collected by the editing author. As his comments are valuable till today and due to current studies on African Campylopoideae by the editing author, this manuscript is more or less unchanged published in its original form.
  • Volume 23 (April 20, 2021)
    Pócs, T. (2021). Notes on the bryophytes of Madagascar 4. Campylopus cuspidatus ssp. frahmii ssp. nov. Frahmia 23:1-7.
    Abstract: A moss at the shore of a subalpine pond on the Marojezy Mountain summit plateau, Madagascar, proved to be a cucullate variant of Campylopus cuspidatus (Hornsch.) Mitt., a species hitherto known only from the Neotropics. As it differs in several features from Campylopus cuspidatus var. dicnemioides (Müll.Hal.) J.-P.Frahm, it is described as ssp. frahmii.
  • Volume 24 (June 07, 2021)
    Sass-Gyarmati, Andrea; Pócs, T. (2021). A new taxon of Bazzania (Lepidoziaceae) collected in Manongarivo Special Reserve, NW Madagascar. Frahmia 24:1-5.
    Abstract: From the rich collections made by Patricia Geissler in Madagascar one taxon of Lepidoziaceae proved to be new to science: Bazzania decrescens (Lehm. et Lindenb.) Trevis. ssp. ambahatrae Gyarmati et Pócs ssp. nov. and here validly published.
  • Volume 25 (August 5, 2021)
    Schäfer-Verwimp, A.; Winter, G.; Yao, K.-Y. (2021). Diobelonella rotundata comb. nov., a new generic record for Taiwan (Aongstroemiaceae, Bryopsida). Frahmia 25:1-10.
    Abstract: Anisothecium rotundatum Broth. (= Dicranella rotundata [Broth.] Takaki) is newly combined under Diobelonella, and the latter genus is reported from Taiwan for the first time. Description of the plants from Taiwan along with photo plates and line drawings of the stem and leaf cross sections of the genera Diobelonella and Dichodontium are provided. Dicranella papua-palustris from Papua New Guinea is considered to be a synonym of Diobelonella rotundata.
ISSN: 2199-4897

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